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"Although Chuck headed an experimental lab throughout much of his career, he was really a theoretical neuroscientist at heart, before theoretical neuroscience was a respected and well-defined subfield. His PhD dissertation, ‘A quantitative theory of neural interactions: theoretical and experimental investigations’, lays out a mathematical theory of computation in the nervous system. Strikingly, it anticipates many subsequent debates — some still raging today, almost 60 years later — about rate versus temporal coding and population coding. Indeed, the thesis includes several chapters and appendices devoted to mathematical derivations and only a relatively small number of experimental figures."
