Not the point of the paper, but this intro bit was news to me:

"Indeed, the host immunity-boosting phenotype may explain the initial spread of Wolbachia in previously uninfected populations. In nematodes, elimination of Wolbachia induces host sterility, and antibiotic treatment is an effective addition to pharmacological treatment of human-infecting, #Wolbachia-positive filarial nematodes"

https://journals.plos.org/plosbiology/article?id=10.1371/journal.pbio.3001972