'Therefore, to predict the dynamics of adaptive radiation, it is necessary to describe not just the individual fitness landscape but also the surface of mean phenotypes and mean fitness: that is, the adaptive landscape. Accomplishing these tasks is such a tall order that a formal adaptive landscape has never been estimated for an adaptive radiation in its natural environment.'

https://academic.oup.com/evolut/article/77/12/2533/7261081?searchresult=1